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A cell wall is not an impenetrable fortress around the delicate plant cell. There are small holes, called plasmodesmata, in the cell walls between plant cells. The cell membranes of neighboring cells are able to connect through these holes. The connections allow the transfer of nutrients, waste, and ions (symplastic pathways). Molecules can also pass through the spaces within the cell walls, avoiding the cells completely (apoplastic pathways).  

It is great that nutrients can move from cell to cell, but there is also a problem with all the holes. Cells can lose water. Plants lose large amounts of water in the middle of the day or on very hot days. When the air heats up and the water vapor pressure decreases, plants lose water through the process of transpiration. The water escapes through pores on the surface of the plant called stomata. Even when the plant cells lose water, the basic shape is maintained by the cell walls. The plant may droop or wilt, but it can recover when water returns to the system. It will look just the same as when it started.  

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