in the artical "How modern life deplets our gut microbes" what organisms would be considered the "keystone"species that the artical is refuring to ? Someone please help ASAP

Peat deposits are found in waterlogged places where dead plant material piles up. when peat is buried, it undergoes compaction. it is subjected to heat and pressure for a long time, and a lot of water is squeezed out during this process. nearly ten vertical feet of original peat material becomes one vertical foot of coal. which of these remains unchanged in the system during the conversion of peat to coal? a) area b) density c) mass d) volume