If the beetles survive by consuming cycad pollen, then whether the beetles should be considered mutualists with, or parasites of, the cycads depends upon A) the extent to which their overall activities affect cycad reproduction. B) the extent to which the beetles are affected by the neurotoxins. C) the extent to which the beetles damage the cycad flowers. D) the distance the beetles must travel between cycad microsporophylls and cycad megasporophylls